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Rory Best will captain Ireland from Wales on his final home appearance in Dublin on Saturday.
Greatest, who will retire after the Word Cup in Japan, leads a side for the closing warm-up match of Ireland.
As anticipated, Johnny Sexton makes his first start at fly-half while Robbie Henshaw and Keith Earls feature from the backline.
Jean Kleyn, who was chosen over Devin Toner at Joe Schmidt squad, begins together with James Ryan at lock.
Schmidt, that will bid farewell to the Aviva Stadium after the Wales match, has generated 12 modifications from last weekend’s victory.
Sexton, Henshaw and Earls make their appearances of their warm-up fittings while Jordan Larmour replaces Jacob Stockdale on the wing.
Will Addison, that had been omitted in the World Cup group, in full-back is come in for by rob Kearney.
There’ll be focus on Sexton venture with Conor Murray following a disappointing Six Nations campaign.
It is a completely new front row in the Wales success while Kleyn has been given the nod ahead of Ulster’s Iain Henderson to begin beside James Ryan at lock as Tadhg Furlong and Cian Healy join Best.
While Josh van der Flier and CJ Stander come at flanker for Peter O’Mahony and Tadhg Beirne jack Conan retains his place at the base of the scrum.
Best says that his final appearance at the Aviva”isn’t something I have given a lot of thought about” but admits that the emotion could strike him after the game.
“I believe for me it’s about making sure we focus on our final game before we board the plane,” stated the 37-year-old.
“When it comes to the final whistle the realisation will set in but it’s important for me and the team it doesn’t influence our preparation.
“I am sure It’ll be a very big occasion once I return on it, but as a team we must execute and require a enormous step forward.”
While there’s been much talk about Schmidt’s 31-man World Cup squad’s announcement feels that it has brought the group closer together,
“It is funny how it looks like there’s so much more room,” he added.
“You get an additional spring in your step and it’s a fantastic time to be on the group.
“Before that, the blade is hanging around you mind regardless of where you’re in the purchase.
“That creates drive and today we are in a 31 it is about us becoming our groups and sharing every little detail, and that’s what makes a really good squad.”
Defence coach Andy Farrell feels that the players have attracted added energy to the group and is now called to raise the tempo.
“They have been held back for a couple weeks and are raring to go,” said Farrell on the debut of Sexton, Henshaw and Earls.
“It is important that we put into a cohesive operation and possess the perfect mental attitude heading into the World Cup.
“It is about trying a number of distinct mixes too. The group is down to 31 gamers but there’s still massive competition for positions.
“It’s our final game before we get on the airplane on Wednesday and we are following a performance that we’re proud of.”
Ireland: Kearney; Larmour, Henshaw, Aki, Earls; Sexton, Murray; Healy, Best (capt), Furlong, Ryan, Kleyn, Stander, van der Flier, Conan.
Replacements: Cronin Porter Ruddock, McGrath, Carty, Ringrose
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Johnny Sexton is now called on Ireland to keep because they last World Cup trainings from Wales, building.
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